I think I figured out what's wrong with my thumb joints: the ligaments at the base of my thumbs are slightly sprained or strained from harmonics. I have been neglecting my hand strengthening exercises, so yes, my muscles were weakened and allowed my thumbs to twist too much when plucking the harmonics in the two pieces I'm working on. And I played an exercise for kicks that was nothing but harmonics. So yeah. Need to stop that for a while. I'm relieved, though, that they've been hurting long enough that it's time to apply heat - as opposed to the immediate 24-48 hr post injury icing that I skipped - because warm hands are such a luxury for me! And I've been doing my exercises again and they seem to be helping.
Sources:
http://black-rose227.deviantart.com/art/Anatomy-of-the-Hand-and-Wrist-270462040
http://www.overtimepainrelief.com/pain-treatment/arthritis-joint-pain/ligament-pain/
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